The story of this Anime starts in a small underground village, where a young boy is working hard to earn his daily meal. His job: Dig deeper into the ground to allow the village to expand. One day, several unexpected things happen. He unearths a large face belonging to a strange mecha. just as he's about to show it to his best friend/older brother, the ceiling collapses and a gigantic metal monstrosity fall through, followed by a pair of boobs, attached to a sniper rifle of immense proportions, and a redhead. A smart little critter joins them and the four of them of them use the recently unearthed mecha to make their escape and rise to the surface. Acquiring a second (much larger) mecha, they set out, on an adventure of epic proportions, with one goal: to go where no man has gone before, to kick the butt of anyone trying to oppose them, and to look cool while doing so. On their journey, they are joined by a VERY gay engineer, a young man and two children from another village, as well as a second group of daring adventurers, and eventually an entire army of people inspired by their sheer awesomeness. Together, they overthrow the planet's tyrant, and his army of beastmen, and pave a way for all of humanity, at the cost of a great sacrifice, though.
The boy who was once a simple digger, now became a hero, and leader of all mankind. The empire he built, however attracts the attention of an enemy unlike anything he ever faced before, threatening to destroy all he struggled to build, and turning his friends into enemies. Unphased by his misfortune, he stands stout, and when faith conspires against him, he punches it in the face. Regaining the trust of his friends, and earning the respect of his former enemies, He sets out to fight one last battle, and succeed where everyone else has failed. Standing over all creation, he will shout into the void: "WHO THE HELL DO YOU THINK I AM!?" as he changes the fate of the entire universe.

Humongous mecha within humongous mecha. An epic battle that will shake the very foundations of the universe. That is Tengen Toppa Gurren Lagann, another masterpiece of Studio Gainax. Oh. It also has a cute animal.